Epigenetics (GENE206 / PATH206)

The topic of the course is epigenetics, the mechanisms by which phenotypes not determined by the DNA sequence itself are nevertheless stably inherited in successive cell divisions.  The course focuses on weekly discussions of primary research papers in the field.  These papers are supplemented with appropriate review articles to provide context and background.  We will begin with the discovery of position-effect variegation in Drosophila in the 1920s and proceed to present day studies of covalent modifications of histones and DNA methylation.  Topics to be covered include position effect, gene silencing, heterochromatin, centromere identity, genomic imprinting, the “histone code”, variant histones, and the role of epigenetics in cancer.  The class will meet once a week for two hours in a discussion format.  The course is designed for graduate students in the Biosciences.

Prerequisite: Successful completion of GENE203 or permission of instructor.