Personal Information:
  Born: March 13, 1946 in Paterson, New Jersey
  Name: March 13, 1998, changed to Joan

Education:
  1971: M.S., Ph.D., Biology, Harvard University
  1968: A.B., Philosophy, University of Rochester, with Highest Honors
  1968: B.S., Biology, University of Rochester, with Distinction and Phi Beta Kappa

Academic Positions:
  1972-present: Stanford University
    Professor of Biological Sciences and of Geophysics
  1970-1972: University of Massachusetts
   Assistant Professor and Instructor of Biology

Professional Experience and Recognition:
  1998: Center Fellow, National Center for Ecological Synthesis and Analysis, UC Santa Barbara
  1995: Dinkelspiel Award for Undergraduate Teaching, Stanford University
  1994: Visiting Research Fellow, Merton College, University of Oxford
  1993: Fellow, American Academy of Arts and Sciences
  1985: Fellow, Guggenheim Foundation
  1978: University Fellow, Stanford University
  Director and Founder, Earth Systems Program, Stanford University, 1992-99
  National Workshops: Theory of Ecological Economics, Santa Barbara, CA
   (1996), Ecology for a Changing Earth, Santa Fe, NM. (1988), and
   Perspectives in Theoretical Ecology, Asilomar, CA. (1987)
  Editorial Boards: American Naturalist (1984-89), Oecologia (1979-82),
   and Theoretical Population Biology (1975-86)
  National Grant Review Panels: NSF (General Ecology, 1987), and DOE (1988)
  National Oceanographic Steering Committees: GLOBEC (1990-91), CoOP (1992-93)
  Science Advisory Boards: Pacific Ocean Conservation Network, California (1997-99)
   Channel Islands National Marine Sanctuary, Santa Barbara (1999-01)
   EPA Science Advisory Board Committee on Valuing the Protection of Ecological Systems
      and Services (2003-present)
  Nonprofit Organization Board: Oceanic Society, San Francisco, (2002-present)
  Community Service: Neighborhood Parks Council; San Francisco Committee on Parks, Recreation,
   and Open Space; Candidate for Supervisor, District 6, San Francisco. (2000-present.)

Postdoctoral Fellows (Primary Advisor)

Dr. Joan Herbers, 1978-1979  Dr. David King, 1979-1981
Dr. Yoh Iwasa, 1981-1983  Dr. Elizabeth Denley, 1982-1984
Dr. Steven Gaines, 1982-1987  Dr. Stephen Brown, 1984-1986
Dr. Sheryl Horowitz, 1985-1986  Dr. Hugh Possingham, 1987-1988
Dr. Terrence Farrell, 1987-1989  Dr. Lloyd Goldwasser, 1988-1990
Dr. Douglas Stoner, 1989-1991  Dr. Timothy Pennington, 1989-1991
Dr. Yehoshua Shkedy, 1992-1995  Dr. Jef Huisman, 1997-1998

Ph.D. Students (Primary Advisor)

Dr. Ronald Yoshiyama, 1972-1977  Dr. David Heckel, 1975-1980
Dr. Stephen Pacala, 1978-1982  Dr. John D. Rummel, 1979-1984
Dr. Kenneth Naganuma, 1982-1989  Dr. Sally Blower, 1982-1987
Dr. John McLaughlin, 1984-1990  Dr. Kristina Miller, 1987-1992
Dr. Susan Alexander, 1988-1993  Dr. Roman Dial, 1988-1992
Dr. Brian Grantham, 1991-1997  Dr. Sharoni Shafir, 1991-1995
Dr. Eric Bjorkstedt, 1992-1997  Dr. Sean Connolly, 1994-1998
Dr. Henrique Pereira, 1997-2002  Dr. Paul Armsworth, 1999-2003
Dr. Lauren Buckley, 2001-2005  Mr. Erol Akcay, 2004-present

Books

2006, Roughgarden, J., Evolution and Christian Faith: Reflections of an Evolutionary Biologist. Island Press. Washington DC. in press.

2004, Roughgarden, J., Evolution's Rainbow: Diversity, Gender, and Sexuality in Nature and People. University of California Press. Berkeley CA. 474 pp. Awarded 2005 Stonewall Prize for nonfiction from American Library Association.

1998, Roughgarden, J., Primer of Ecological Theory. Prentice Hall, Upper Saddle River, New Jersey. 450 pp.

1995, Roughgarden, J., Anolis Lizards of the Caribbean: Ecology, Evolution, and Plate Tectonics. Oxford University Press, 200 pp., illustr..

1989, Roughgarden, J., R. May, and S. Levin, Eds.,   Perspectives in Ecological Theory. Princeton University Press, Princeton, New Jersey, vii + 394 pp., illustr.

1987, Ehrlich, P., and J. Roughgarden, The Science of Ecology. Macmillan Publishing Company, New York, New York, xxv + 710 pp., illustr.

1979, Roughgarden, J., Theory of Population Genetics and Evolutionary Ecology: An Introduction. Macmillan Publishing Company, New York, 612 pp., illustr. (Reprinted 1987 by Macmillan, and in 1996 by Prentice Hall).

Selected Articles

2006, Roughgarden J., M. Oishi, and E. Akcay. Reproductive social behavior: Cooperative games to replace sexual selection. Science. In press.

2006, Roughgarden J. Foreward. Pp. xvii--xx in: Marine Metapopulations J. Kritzer and P. Sale (eds.), Elsevier, Burlington, MA. In press.

2006, Buckley, L. and J. Roughgarden. A hump-shaped density-area relationship for island lizards. Oikos In press.

2005, Roughgarden. J. The myth of sexual selection. California Wild 53(3):18--23

2005, Buckley, L. and J. Roughgarden. Lizard habitat partitioning on islands: the interaction of local and landscape scales. Journal of Biogeography 32(12): 2113--2121.

2005, Buckley, L. and J. Roughgarden. Effect of species interactions on landscape abundance patterns. Journal of Animal Ecology 74:1182--1194.

2005, Armsworth, P. and J. Roughgarden. Disturbance induces the contrasting evolution of reinforcement and dispersiveness in directed and random movers. Evolution 59(10):2083--2096.

2005, Armsworth, P. and J. Roughgarden. The impact of directed versus random movement on population dynamics and biodiversity patterns. American Naturalist 165:449-465.

2004, Pereira, H., G. Daily, and J. Roughgarden. A framework for assessing the relative vulnerability of species to land-use change. Ecological Applications. 14(3):730--742.

2004, Roughgarden, J. Twist in the tale of two genders. Times Higher Education Supplement. June 4, 2004. 1643:20-21.

2004, Roughgarden, J. The in-crowd. New Scientist 181:36-39.

2004, Roughgarden, J. Evolution and the embodiment of gender. GLQ: A Journal of Lesbian and Gay Studies. 10:287-291.

2004, Gerber, L., O. Reichman, and J. Roughgarden. Food hoarding: Future value in optimal foraging decisions. Ecological Modelling. 175:77-85.

2003, Hughes, T., A. Baird, D. Bellwood, M. Card, S. Connolly, C. Folke, R. Grosberg, O. Hoegh-Guldberg, J. Jackson, J. Kleypas, J. Lough, P. Marshall, M. Nyström, S. Palumbi, J. Pandolfi, B. Rosen, and J. Roughgarden. Climate change, human impacts, and the resilience of coral reefs. Science 301:929-933.

2003, Armsworth, P. and J. Roughgarden. The economic value of ecological stability. Proc. Nat. Acad. (USA) 100:7147-7151.

2003, Pereira, H., A. Bergman, and J. Roughgarden. Socially stable territories: The negotiation of space by interacting foragers. American Naturalist 161:143-152.

2002, Bjorkstedt, E., L. Rosenfeld, B. Grantham, Y.  Shkedy, and J. Roughgarden. Distributions of larval rockfish (Sebastes spp.) across nearshore fronts in a coastal upwelling region. Marine Ecology Progress Series 242:215-228.

2002, Balmford, A., A. Bruner, P. Cooper, R. Costanza, S. Farber, R. Green, M. Jenkins, P. Jefferiss, V. Jessamy, J. Madden, K. Munro, N. Myers, S. Naeem, J. Paavola, M. Rayment, S. Rosendo, J. Roughgarden, K. Trumper, and K. Turner. Economic reasons for conserving wild nature. Science 297:950-953.

2002, Pereira, H., S. Loarie, and J. Roughgarden. Monogamy, polygyny and interspecific interactions in the lizards Anolis gingivinus and Anolis pogus. Caribbean Journal of Science, 38:8-12.

2002, Bascompte, J. H. Possingham, and J. Roughgarden. Patchy populations in stochastic environments: Critical number of patches for persistence. Amer. Natur. 159:128-137.

2001, Connolly, S., B. A. Menge and J. Roughgarden. A latitudinal gradient in recruitment of intertidal invertebrates in the Northeast Pacific Ocean. Ecology, 82:1799-1813.

2001, Roughgarden, J. and P. Armsworth. Managing ecosystem services. In: Press, M. N. Huntly, and S. Levin, (eds.) Ecology: Achievement and Challenge, Blackwell Science, pp. 337-356.

2001, P. Armsworth and J. Roughgarden. An invitation to ecological economics, Trends in Ecology and Evolution 16:229-234.

2001, Roughgarden, J. Guide to diplomatic relations with economists. Bull. Ecol. Soc. America 82:85-88.

2000, Hughes, J. and J. Roughgarden. Species diversity and biomass stability. American Naturalist, 155:618-627.

1999, Connolly, S. and J. Roughgarden. Theory of marine communities: competition, predation and recruitment-dependent interaction strength. Ecological Monographs, 69:277-296.

1999, Connolly, S. and J. Roughgarden. Increased recruitment of northeast Pacific barnacles during the 1997 El Niño. Limnol. Oceanogr. 44:466-469.

1999, S. Murray, R. F. Ambrose, J. A. Bohnsack, L. W. Botsford, M. H. Carr, G. E. Davis, P. K. Dayton, D. Gotshall, D. R. Gunderson, M. A. Hixon, J. Lubchenco, M. Mangel, A. MacCall, D. A. McArdle, J. C. Ogden, J. Roughgarden, R. M. Starr, M. J. Tegner, and M. M. Yoklavich. No-take reserve networks: sustaining fishery populations and marine ecosystems. Fisheries Nov. 1999 pp. 11-25.

1998, Connolly, S. and J. Roughgarden. A range extension for the volcano barnacle, Tetraclita rubescens. California Fish and Game 84:182-183.

1998, Hughes, J. and J. Roughgarden. Aggregate community properties and the strength of species' interactions. Proc. Nat. Acad. Sci. (USA), 95:6837-6842.

1998, Dial, R., and J. Roughgarden. Theory of marine communities: the intermediate disturbance hypothesis, Ecology 79:1412-1424.

1998, Shafir, S. and J. Roughgarden. Testing predictions of foraging theory for a sit-and-wait forager, Anolis gingivinus. Behavioral Ecology 9:74-84.

1998, Connolly, S. and J. Roughgarden, A latitudinal gradient in northeast Pacific intertidal community structure: evidence for an oceanographically based synthesis of marine community theory. American Naturalist, 151:311-326.

1998, Roughgarden, J., How to manage fisheries. Ecological Applications, 8(1):S160-S164.

1998, Roughgarden, J., Production functions from ecological populations: a survey with emphasis on spatially explicit models. In: Tilman, D. and P. Kareiva, (eds.) Spatial Ecology: The Role of Space in Population Dynamics and Interspecific Interactions, Princeton University Press, pp. 296-317.

1998, Boggs, C. and J. Roughgarden. Biodiversity: result of speciation and extinction. Chapter 17 in G. Ernst, ed., Earth Systems: Processes and Issues, Cambridge University Press, in press.

1998, Boggs, C. and J. Roughgarden. Evolution: adaptation and environmental change. Chapter 19 in G. Ernst, ed., Earth Systems: Processes and Issues, Cambridge University Press, in press.

1997, Shkedy, Y., and J. Roughgarden, Barnacle recruitment and population dynamics predicted from coastal upwelling. Oikos 80(3): 487-498.

1997, Bjorkstedt E., and J. Roughgarden, Larval transport and coastal upwelling: an application of HF radar in ecological research. Oceanography 10(2): 64-67.

1997, Brown, G. and J. Roughgarden, A metapopulation model with private property and a common pool. Ecological Economics, 22:65-71.

1997, Goldwasser, L. and J. Roughgarden, Sampling effects and the estimation of food web properties. Ecology, 78:41-54.

1996, Dial, R., and J. Roughgarden, Natural history observations of Anolisomyia rufianalis (Diptera: Sarcophagidae) infesting Anolis lizards in a rain forest canopy. Environ. Entomol. 25:13:25-1328.

1996, Alexander, S.E. and J. Roughgarden, Larval transport and population dynamic of intertidal barnacles: a coupled benthic/oceanic model. Ecological Monographs, 66:259-275.

1996, Roughgarden, J. and F. Smith, Why fisheries collapse and what to do about it. Proc. Nat. Acad. Sci., (USA), 93:5078-5083

1996, Shafir, S. and J. Roughgarden, The effect of memory length on the individual fitness of a lizard. In Belew, R.K. and M. Mitchell (Eds.), Adaptive Individuals in Evolving Populations: Models and Algorithms, Santa Fe Institute Studies in the Science of Complexity, Addison Wesley, Reading, MA., pp. 173-182.

1996, Roughgarden, J., A. Bergman, S. Shafir and C. Taylor, Adaptive computation in ecology and evolution: a guide for future research. In Belew, R.K. and M. Mitchell (Eds.), Adapting Individuals in Evolving Populations: Models and Algorithms, Sante Fe Institute Studies in the Science of Complexity, Addison Wesley, Reading, MA., pp. 25-32.

1996, Roughgarden, J., Ecology. In: Encyclopedia of Climate and Weather.

1995, Rosenfeld, L., T. Anderson, G. Hatcher, J. Roughgarden, and Y. Shkedy, Upwelling fronts and barnacle recruitment in central California. Technical Report 95-19, Monterey Bay Aquarium Research Institute. Moss Landing, CA

1995, Roughgarden, J., Can economics protect biodiversity? Pp. 149-156 in: T. Swanson, Ed., The economics and ecology of biodiversity decline. Cambridge University Press.

1995, Holling, C.S., D.W. Schindler, B.W. Walker and J. Roughgarden, Biodiversity in the functioning of ecosystems: an ecological synthesis. In Perings, C., C.G. Maler, C. Folke, C.S. Holling, and B.O. Jansson (Eds.), Biodiversity Loss: Economic and Ecological Issues, Cambridge, Cambridge University Press, pp. 44-83.

1995, Brown, G. and J. Roughgarden, An ecological economy: notes on harvest and growth. In Perrings, C., K.G. Maler, C. Folke, C.S. Holling and B.O. Jansson (Eds.), Biodiversity Loss: Ecological and Economic Issues, Cambridge, Cambridge University Press, pp. 150-189. (Reprinted in: J. Barkley Rosser, Jr., 2004. Complexity in Economics, Volume III. The International Library of Critical Writings in Economics 174. Edward Elgar, Cheltenham, UK, pp. 383-423.)

1995, Roughgarden, J., Vertebrate patterns on islands. In Vitousek, P.M., L.L. Loope, and H. Andersen (Eds.), Islands: Biological Diversity and Ecosystem Function, Springer, Germany, pp. 51-56.

1995, Dial, R., and J. Roughgarden, Experimental removal of insectivores from rain forest canopy: direct and indirect effects. Ecology, 76 (6) pp. 1821-1834.

1994, Shafir, S., and J. Roughgarden, The effect of memory length on the foraging behavior of a lizard. In Cliff, D., P. Husbands, J.A. Meyer, and S.W. Wilson (Eds.), From Animals to Animats 3: Proceedings of the Third International Conference on Simulation of Adaptive Behavior, MIT Press, Cambridge, MA. pp. 221-225.

1994, Shafir, S., and J. Roughgarden, Instrumental discrimination conditioning of Anolis cristatellus in the field with food as a reward. Caribbean Journal of Science, 30 (3-4): 228-233.

1994, Dial, R. and J. Roughgarden, Notes on the absolute abundance of canopy anoles: Anolis cuvieri, Anolis stratulus, and Anolis evermanni (Lacertilia: polychridea) in the Luquillo Forest, Puerto Rico. Caribbean Journal of Science, 30 (3-4): 278-279.

1993, Fernandez, D., C. Teague, J. Vesecky, and J. Roughgarden, Upwelling and relaxation events inferred from high-frequency radar measurements of coastal surface currents. Proceedings of the IGARSS '93 Conference - IEEE International Geoscience and Remote Sensing Symposium, 4:1931-1933.

1993, Miller, K., and J. Roughgarden, Descriptions of the larvae of Tetraclita rubescens and Megabalanus californicus with a comparison of the common barnacle larvae of the central California coast. Journal of Crustacean Biology, pp. 579-600.

1993, Shkedy, Y., D. Fernandez, C. Teague, J. Vesecky, and J. Roughgarden, Detecting upwelling along the central coast of California during an El-Niño year using HF-radar. Continental Shelf Research, 15 (7): 803-814.

1993, Roughgarden, J., H. Possingham, S. Tuljapurker and M. Wilks, Population cycling in space-limited organisms subject to density-dependent predation. The American Naturalist, 143 (4): 563-582.

1993, McLaughlin, J., and J. Roughgarden, Species interactions in space. Chapter 8 in Species Diversity in Ecological Communities: Historical and Geographical Perspectives, Ed. by R. E. Ricklefs and D. Schluter, pp. 89-98.

1993, Roughgarden, J., Pennington and S. Alexander, Dynamics of the rocky intertidal zone with remarks on generalization in ecology. Philosophical Transactions of the Royal Society London B., 341:449-454.

1993, Goldwasser, L., and J. Roughgarden, Construction and analysis of a large Caribbean food web. Ecology, 74:4, pp. 1216-1233.

1992, Dobson, A., S. Pacala, J. Roughgarden, E. Carper, and E. Harris., The Parasites of Anolis lizards in the northern Lesser Antilles. Oecologia, 91:1, pp. 110-117.

1992, Roughgarden, J., Comments on the paper by Losos: Character displacement versus taxon loop. Copeia, 2:288-295.

1992, Koza, J., J. Rice, and J. Roughgarden., Evolution of food-foraging strategies for the Caribbean Anolis lizard using genetic programming. Adaptive Behavior, 1:2, pp. 171-199.

1991, Roughgarden, J., S. Running, and P. Matson, What does remote sensing do for ecology? Ecology, 72:6, pp. 1918-1922.

1991, Miller, K., P. Jones, and J. Roughgarden, Monoclonal antibodies as species-specific probes in oceanographic research: Examples with intertidal barnacle larvae. Molecular Marine Biology & Biotechnology, 1:35-47.

1991, McLaughlin, J., and J. Roughgarden, Predation across spatial scales in heterogeneous environments. Theoretical Population Biology, 40:2, pp. 149-172.

1991, Farrell, T., D. Bracher, and J. Roughgarden, Cross-shelf transport causes recruitment to intertidal populations in central California. Limology & Oceanography, 36:279-288.

1991, McLaughlin, J., and J. Roughgarden, Pattern and stability in predator communities: How diffusion in spatially variable environments affects the Lotka-Volterra model. Theoretical Population Biology, 40:148-172.

1991, Roughgarden, J., J. Pennington, D. Stoner, S. Alexander, and K. Miller, Collisions of upwelling fronts with the intertidal zone: The cause of recruitment pulses in barnacle populations of central California. Acta Oecologica, 12:35-51.

1991, Roughgarden, J., The evolution of sex. American Naturalist, 138(4) pp. 934-985.

1990, Possingham, H., and J. Roughgarden, Spatial population dynamics of an organism with a complex life cycle. Ecology, 71:973-985.

1990, Roughgarden, J., Origin of the Eastern Caribbean: Data from reptiles and amphibians. In D. K. Larue and G. Draper (Eds.), Trans. 12th Caribbean Geological Conference, St. Croix, USVI, Miami Geological Survey, pp. 10-26.

1990, Naganuma, K., and J. Roughgarden, Optimal body size in Lesser Antillean Anolis lizards-A mechanistic approach. Ecological Monographs, 60:239-256.

1990, Brown, J., and J. Roughgarden, Ecology for a changing earth. Bulletin of the Ecological Society of America, 71:173-188.

1989, Roughgarden, J., The structure and assembly of communities. In Roughgarden, R. M. May, and S. A. Levin (Eds.) Perspectives in Ecological Theory, Princeton University Press, pp. 203-226.

1989, Roughgarden, J., Viewpoint: The United States needs an ecological survey. Bioscience, 39:5.

1989, McLaughlin, J., and J. Roughgarden, Avian predation on Anolis lizards in the northeastern Caribbean: An inter-island contrast. Ecology, 70:617-628.

1989, Brown, J., and J. Roughgarden, U. S. ecologists address global change. Trends in Ecology and Evolution, 4:255.

1989, Blower, S., and J. Roughgarden, Parasites detect host spatial pattern and density: A field experimental analysis. Oecologia, 78:138-141.

1989, Miller, K., S. Blower, D. Hedgecock, and J. Roughgarden, Comparison of larval and adult stages of Chthamalus dalli and Chthamalus fissus (Cirripedia: Thoracica). Journal Crustacean Biology, 9:242-256.

1989, Blower, S., and J. Roughgarden, Population dynamics and parasitic castration: A mathematical model. American Naturalist, 134:848-858.

1988, Roughgarden, J., and S. Pacala, Taxon cycle among Anolis lizard populations: Review of evidence. In Daniel Otte and John Endler (Eds.), Speciation and its Consequences, Sinauer Associates, Inc., pp. 403-432.

1988, Blower, S., and J. Roughgarden, Parasitic castration: host species preferences, size-selectivity and spatial heterogeneity. Oecologia, 75:512-515.

1988, Roughgarden, J., S. Gaines, and H. Possingham, Recruitment dynamics in complex life cycles. Science, 241:1460-1466.

1988, Roughgarden, J., The evolution of marine life cycles. In M. W. Feldman (Ed.), Mathematical Evolutionary Theory, Princeton University Press, pp. 270-300.

1987, Roughgarden, J., Community coevolution: A comment. Evolution, 41:5, pp. 1130-1134.

1987, Gaines, S., and J. Roughgarden, Fish in offshore kelp forests affect recruitment to intertidal barnacle populations. Science, 235:479-481.

1987, Roughgarden, J., S. Gaines, and S. Pacala, Supply side ecology: The role of physical transport processes. In J. H. R. Gee and P. S. Giller (Eds.), Organization of Communities, Past and Present, The 27th Symposium of the The British Ecological Society, Blackwell Scientific Publications, pp. 491-518.

1987, Blower, S., and J. Roughgarden, Population dynamics and parasitic castration: Test of a model. American Naturalist, 134:848-858

1986, Roughgarden, J., and Y. Iwasa, Dynamics of a metapopulation with space-limited subpopulations. Theoretical Population Biology, 29:235-261.

1986, Roughgarden, J., and J. Diamond, Overview: The role of species interactions in community ecology. In J. Diamond and T. Case (Eds.), Community Ecology, Harper & Row, New York, pp. 333-344.

1986, Roughgarden, J., A Comparison of food-limited and space-limited animal competition communities. In J. Diamond and T. Case (Eds.), Community Ecology, Harper & Row, New York, pp. 492-516.

1986, Roughgarden, J., Predicting invasions and rates of spread. In Harold Mooney and James A. Drake (Eds.), Ecology of Biological Invasions of North America and Hawaii, Ecological Studies, Springer-Verlag, 58:179-188.

1986, Iwasa, Y., and J. Roughgarden, Interspecific competition among metapopulations with space-limited subpopulations. Theoretical Population Biology, 30:260-214.

1986, Roughgarden, J., T. Schoener, and T. Fenchel, The body-size-prey-size hypothesis: A defense. Ecology, 67:260-261.

1986, Roughgarden, J., Models of population processes for plants. In L. J. Gross and R. M. Miura (Eds.), Some mathematical questions in biology-plant biology. Lectures on Mathematics in the Life Sciences, American Mathematical Society, Providence, R.I., 18:235-267.

1985, Brown, S., and J. Roughgarden, Growth, morphology, and laboratory culture of Balanus glandula larvae ( Cirripedia:Thoracica). J. Crustacean Biology, 5:574-590.

1985, Rummel, J., and J. Roughgarden., Effects of reduced perch-height separation on competition between two Anolis lizards. Ecology, 66:2 pp. 430-444.

1985, Roughgarden, J., Y. Iwasa, and C. Baxter, Demographic theory for an open marine population with space-limited recruitment. Ecology, 66:54-57.

1985, Gaines, S., and J. Roughgarden, Larval settlement rate: A leading determinant of structure in an ecological community of the marine intertidal zone. Proceedings of the National Academy of Sciences, USA, 82:3707-3711.

1985, Gaines, S., S. Brown, and J. Roughgarden, Spatial variation in larval concentrations as a cause of spatial variation in settlement for the barnacle, Balanus glandula. Oecologia, 67:267-272.

1985, Iwasa, Y., and J. Roughgarden, Evolution in a metapopulation with space-limited subpopulation. IMA Journal of Mathematics Applied in Medicine & Biology, 2:93-107.

1985, Pacala, S., and J. Roughgarden, Population experiments with the Anolis lizards of St. Maarten and St. Eustatius. Ecology, 66:129-141.

1985, Rummel, J., and J. Roughgarden, A theory of faunal buildup for competition communities. Evolution, 39:1009-1033.

1985, Holsinger, K., and J. Roughgarden, A model for the dynamics of an annual plant population. Theoretical Population Biology, 28:288-313.

1984, Iwasa, Y., and J. Roughgarden, Shoot/root balance of plants: Optimal growth of a system with many vegetative organs. Theoretical Population Biology, 25:78-105.

1984, Pacala, S., and J. Roughgarden, Control of arthropod abundance by Anolis lizards on St. Eustatius (Neth. Antilles). Oecologia, 64:160-162.

1984, Roughgarden, J., S. Pacala, and J. Rummel, Strong present-day competition between the Anolis lizard populations of St. Maarten (Neth. Antilles). In B. Shorrocks (Ed.), Evolutionary Ecology, The 23rd Symposium of the British Ecological Society, Blackwell Scientific Publications, pp. 203-220.

1984, Roughgarden, J. and N. Chiariello, Storage allocation in seasonal races of an annual plant: Optimal versus actual allocation. Ecology, 65:1290-1301.

1984, Roughgarden, J., S. Gaines, and Y. Iwasa, Dynamics and evolution of marine populations with pelagic larval dispersal. In R. M. May (Ed.), Exploitation of Marine Communities Dahlem Konferenzen, Berlin. Springer-Verlag, pp. 111-128.

1983, Roughgarden, J., D. Heckel, and E. Fuentes, Coevolutionary theory and the biogeography and community structure of Anolis. In R. Huey, E. Pianka and T. Schoener (Eds.), Lizard Ecology: Studies of a Model Organism. Harvard University Press, pp. 371-410.

1983, Roughgarden, J., J. Rummel, and S. Pacala, Experimental evidence of strong present-day competition between Anolis populations of the Anguilla Bank, A preliminary report. In A. Rhodin and K. Miyata, (Eds.), Advances in Herpetology and Evolutionary Biology-Essays in Honor of Ernest Williams. Museum of Comparative Zoology, Harvard University Press, pp. 499-506.

1983, Adolph, S., and J. Roughgarden, Foraging by Passerine birds and Anolis lizards on St. Eustatius (Neth. Antilles): Implications for interclass competition and predation. Oecologia, 56:313-317.

1983, Pacala, S., J. Rummel, & J. Roughgarden, A technique for enclosing Anolis lizard populations under field conditions. J. Herpetology, 17:94-97.

1983, King, D., and J. Roughgarden, Energy allocation patterns of the California grassland annuals, Plantago erecta and Clarkia rubicunda. Ecology, 64:16-24.

1983, Roughgarden, J., The theory of coevolution. In D. Futuyma and M. Slatkin (Eds.), Coevolution, Sinauer Associates, pp. 33-64.

1983, Roughgarden, J., Coevolution between competitors. In D. Futuyma, and M. Slatkin (Eds.), Coevolution, Sinauer Associates, pp. 383-403.

1983, Roughgarden, J., Competition and theory in community ecology. American Naturalist, 122:583-601.

1983, Rummel, J., and J. Roughgarden, Some differences between invasion-structured and coevolution-structured competitive communities - A preliminary theoretical analysis. Oikos, 41:477-486.

1982, Pacala, S., and J. Roughgarden, Spatial heterogeneity and interspecific competition. Theoretical Population Biology, 21:92-113.

1982, King, D., and J. Roughgarden, Multiple switches between vegetative reproductive growth in annual plants. Theoretical Population Biology, 21:194-204.

1982, Shigesada, N., and J. Roughgarden, The role of rapid dispersal in the population dynamics of competition. Theoretical Population Biology, 21:353-372.

1982, King, D., and J. Roughgarden, Graded allocation between vegetative and reproductive growth for annual plants in growing seasons of random length. Theoretical Population Biology, 22:1-16.

1982, Pacala, S., and J. Roughgarden, The evolution of resource partitioning in a multidimensional resource space. Theoretical Population Biology, 22:127-145.

1982, Pacala, S., and J. Roughgarden, Resource partitioning and interspecific competition in two two-species insular Anolis lizard communities. Science, 217:444-446.

1982, Roughgarden, J., W. Porter and D. Heckel, Resource partitioning of space and its relationship to body temperature in Anolis lizard populations. Oecologia, 50:256-264.

1981, Anderson, G., A. Ehrlich, P. Ehrlich, J. Roughgarden, B. Russell, and F. Talbot, The community structure of coral reef fishes. American Naturalist , 117:476-495.

1981, Roughgarden, J., W. Porter, and D. Heckel, Resource partitioning of space and its relationship to body temperature in Anolis lizard populations. Oecologia, 50:256-264.

1980, Heckel, D., and J. Roughgarden, A species near its equilibrium size in a fluctuating environment can evolve a lower intrinsic rate of increase. Proc. Nat. Acad. Sci., 77:7497-7500.

1979, Heckel, D., and J. Roughgarden, A technique for estimating the size of lizard populations. Ecology, 60:966-975.

1979, Roughgarden, J., Coevolution in ecological systems III. Coadaptation and equilibrium population size. In P. F. Brussard (Ed.), Ecological Genetics: The Interface, Springer-Verlag, New York, pp. 27-50.

1979, Roughgarden, J., A local concept of structural homology for ecological communities with examples from simple communities of West Indian Anolis lizards. In U. Halbach and J. Jacobs (Eds.), Population Ecology, Fortschritte der 149-158.

1978, Roughgarden, J., Influence of competition on patchiness in a random environment. Theoretical Population Biology, 14:185-203.

1977, Roughgarden, J., Patchiness in the spatial distribution of a population caused by stochastic fluctuations in resources. Oikos, 29:52-59.

1977, Roughgarden, J., Coevolution in ecological systems: Results from ``loop analysis" for purely density-dependent coevolution. In F. B. Christiansen and T. M. Fenchel (Eds.), Measuring Selection in Natural Populations, Vol. 19 in Lecture Notes in Biomathematics. Springer-Verlag, New York, pp. 499-517.

1977, Roughgarden, J., and E. Fuentes, The environmental determinants of size in solitary populations of West Indian Anolis lizards. Oikos, 29:44-51.

1977, Yoshiyama, R., and J. Roughgarden, Species packing in two dimensions. American Naturalist, 111:107-121.

1976, Roughgarden, J., Resource partitioning among competing species: A coevolutionary approach. Theoretical Population Biology, 9:388-424.

1976, Roughgarden, J., A simple model for population dynamics in stochastic environments. American Naturalist, 109:713-736.

1975, Roughgarden, J., Populations dynamics in a stochastic environment: Spectral theory for the linearized N-species Lotka-Volterra competition equations. Theoretical Population Biology, 7:1-12.

1975, Feldman, M., and J. Roughgarden, A population's stationary distribution and chance of extinction in a stochastic environment with remarks on the theory of species packing. Theoretical Population Biology, 7:197-207.

1975, Roughgarden, J., and M. Feldman, Species packing and predation pressure. Ecology, 56:489-492.

1975, Roughgarden, J., Evolution of marine symbiosis - a simple cost-benefit model. Ecology, 56:1201-1208.

1974, Roughgarden, J., The fundamental and realized niche of a solitary population. American Naturalist, 108:232-235.

1974, Roughgarden, J., Species packing and the competition function with illustrations from coral reef fish. Theoretical Population Biology, 5:163-186.

1974, Roughgarden, J., Niche width: Biogeographic patterns among Anolis lizard populations. American Naturalist, 108:429-442.

1974, Roughgarden, J., Population dynamics in a spatially varying environment: How population size ``tracks" spatial variation in carrying capacity. American Naturalist, 108:649-664.

1972, Roughgarden, J., Evolution of niche width American Naturalist, 106:683-718.

1971, Roughgarden, J., Density-dependent natural selection Ecology, 52:453-468.